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Mechanisms of Action & Metabolic Signaling Pathways MOTS-c operates through novel nuclear-mitochondrial communication cascades: ┌──► AMPK Phosphorylation (Triggers Cellular Energy Sensing) │ MOTS-c Micropeptide ┼──► Nuclear Translocation (Modulates Stress-Response Gene Expression) │ └──► Folate-Purine Synthesis Regulation & Glucose Transporter (GLUT4) Upregulation 1