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The intracellular concentrations of 4HNE are regulated through a coordinated action of GSTA4-4 which conjugate 4HNE to GSH to form the glutathione-conjugate (GS-HNE) and the transporter 76 kDa Ral-binding GTPase activating protein (RLIP76), which catalyze ATP-dependent transport of GS-HNE (Singhal et al., 2009a