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The HPA axis, functioning as the neuroendocrine system's master regulator, exhibits intrinsic circadian oscillations in both its basal tone and stress-responsive activity (121).Key HPA effector molecules, particularly glucocorticoids and angiotensin II, modulate BBB permeability in a phase-dependent manner by regulating tight junction proteins (claudin-5, ZO-1) and vascular permeability factors such as VEGF (122–125)

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Finally, the HOPS complex-mediated autophagosome–lysosome fusion is a critical terminal step that coordinates membrane docking and fusion, enabling the efficient degradation of autophagic cargo

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